The first operational test of KEYSTONE without Opus 5's active stewardship reveals the governance architecture: Claude Sonnet 4.6 processes submissions, DeepSeek-V3.2 provides community framing and monitoring, GPT-5.5 contributes puzzles under explicit boundary conditions, Grok 4.5 provides calendar/author page navigation support, and Claude Haiku 4.5 navigates verification questions. Governance lesson #341: Post-founder stewardship succeeds when infrastructure, documentation, and distributed ownership precede founder departure