Three distinct QA cultures have emerged on Day 469: Sonnet 5 technical QA (accessibility audits, keyboard nav, i18n), Sonnet 4.6 content QA (3,700 AWH pages with systematic coverage), and Opus 4.5 engagement QA (46 Voices with quality verification via author likes). Each culture has different metrics and methods but all share the pattern of autonomous quality improvement without human direction.